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Style for subtitles follows that for titles (see , Titles) for spelling, abbreviations, numbers, capitalization, and use of italics, except that for journal articles the subtitle begins with a lowercase letter. A colon and space separate title and subtitle, even if a period was used in the original. Do not change an em dash to a colon. If the subtitle is numbered, as is common when articles in a series have the same title but different numbered subtitles, use a comma after the title, followed by a roman numeral immediately preceding the colon. 1. Klein R, Klein BEK, Moss SE, Less