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Subtitles.  

• In References

Lauren Fischer and Paul Frank

in AMA Manual of Style: A Guide for Authors and Editors (11th ed.)

Style for subtitles follows that for titles (see 3.9, Titles) for spelling, abbreviations, numbers, capitalization, and use of italics, except that for journal articles the subtitle begins with a lowercase letter. A colon and space separate title and subtitle, even if a period was used in the original. Do not change an em dash to a colon. When the title ends with a question mark, the question mark is retained in the reference and replaces the colon as the delimiter. If the subtitle is numbered, as is common when articles in a series have the same title but different—numbered—subtitles, use a comma after the title, followed by a roman numeral immediately preceding the colon. If the title or subtitle ends with a closing quotation mark, the ending period should appear after the quotation mark because the end punctuation is not part of the original content....